Optimus, damaged in Mexico City and disguised as an old Marmon 97 semi-truck, is discovered in an abandoned theater by Cade Yeager, a financially struggling Texan inventor, and brings him back to his farm. While his teenage daughter Tessa and business partner Lucas Flannery encourage him to turn Optimus over to the authorities after realizing what he really is, Cade instead fixes Optimus, hoping to understand his technology and unexpectedly revives him. Hoping to collect a reward for turning Optimus in to the government, Lucas calls the authorities, which gets redirected to Cemetery Wind, who, led by Lockdown and Attinger's second-in-command James Savoy (who hates Transformers because he lost his sister in Chicago), attacks and destroys the farm, but Optimus and Tessa's boyfriend, Irish rally car driver Shane Dyson, come to the family's aid. While escaping Cemetery Wind and Lockdown, Lucas is killed by one of Lockdown's grenades. Out in the desert, they seek refuge in an abandoned gas station. Later, Optimus gains an alternate form after scanning a passing Western Star 5700 XE Phantom Custom truck with the original red and blue paint with flames (healing his injuries in the process) and summons the surviving Autobots – Bumblebee, who has assumed a new alternate form (a 1967 Chevrolet Camaro), Hound, Drift, and Crosshairs – who have come to distrust humans due to Cemetery Wind constantly hunting them down. Using a CIA drone which he stole during the home invasion, Cade discovers K. S. I. 's involvement in the attacks on the Autobots. Optimus vows to personally kill Attinger for his actions against his brethren.
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